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All current publications: see Google Scholar profile

Publications through 2015: PDFs are linked below


(in press). Schriever, T.A., M.T. Bogan, K.S. Boersma, M. Cañedo-Argüelles, K.L. Jaeger, J.D. Olden, & D.A. Lytle.  Hydrology shapes taxonomic and functional structure of desert stream invertebrate communities. Freshwater Science.

Phillipsen, I.V., E.H. Kirk, M.T. Bogan, M.C. Mims, J.D. Olden, D.A. Lytle. Dispersal ability and habitat requirements determine landscape-level genetic patterns in desert stream insects. Molecular Ecology 24(1): 54-69.

(in press). Mims, M.C. I.C. Phillipsen, D.A. Lytle, E.H. Kirk, & J.D. Olden. Ecological strategies predict associations between aquatic and genetic connectivity for dryland amphibians. Ecology.

Cañedo-Argüelles, M., K.S. Boersma, M.T. Bogan, J.D. Olden, I.C. Phillipsen, T.A. Schriever, D.A. Lytle. Dispersal strength determines metacommunity structure in a dendritic riverine network. Journal of Biogeography 42(4): 778-790.

(in press). Lytle, D.A. Aquatic Hemiptera. Pp. in Thorp, J.H., D.C. Rogers, & K. Tockner (Eds.): Thorp and Covich's Freshwater Invertebrates, Fourth Edition. Elsevier Press.


(in press). Bogan, M.T., K.S. Boersma, & D.A. Lytle. Resistance and resilience to seasonal and supraseasonal drought in invertebrate communities of arid-land headwater streams. Freshwater Biology.

(in press). Boersma, K.S., & D.A. Lytle. Overland dispersal and drought escape behavior in a flightless aquatic insect, Abedus herberti (Hemiptera: Belostomatidae). Southwestern Naturalist.

Olden, J.D., C.P. Konrad, T.S. Melis, M.J. Kennard, M.C. Freeman, M.C. Mims, E.N. Bray, K.B. Gido, N.P. Hemphill, D.A. Lytle, L.E. McMullen, M. Pyron, C.T. Robinson, J.C. Schmidt, & J.G. Williams. Are large-scale flow experiments informing the science and management of freshwater ecosystems? Frontiers in Ecology and the Environment 12: 176-185. doi:10.1890/130076.

Boersma, K.S., M.T. Bogan, B.A. Henrichs, & D.A. Lytle. Top predator removals have consistent effects on large species despite high environmental variability. Oikos. DOI: 10.1111/oik.00925

Boersma, K.S., M.T. Bogan, B.A. Henrichs, & D.A. Lytle. Invertebrate assemblages of pools in arid-land streams have high functional redundancy and are resistant to severe drying. Freshwater Biology 59: 491-501.

Bogan, M.T., O.G. Gutiérrez, J.A. Alvarado, & D.A. Lytle. New records of Martarega, Graptocorixa, and Abedus (Heteroptera: Notonectidae, Corixidae, Belostomatidae) from northwestern Mexico and Arizona, including the first record of Graptocorixa emburyi in the United States. Southwestern Naturalist 58(4) 494-497.

Bogan, M.T., N. Noriega-Felix, S. Leonor Vidal-Aguilar, L.T. Findley, D.A. Lytle, O.G. Gutiérrez, J.A. Alvarado, and A. Varela-Romero. Biogeography and conservation of aquatic fauna in spring-fed tropical canyons of the southern Sonoran Desert, Mexico. Biodiversity & ConservationDOI 10.1007/s10531-014-0745-z


Phillipsen, I.C., & D.A. Lytle. Aquatic insects in a sea of desert: Population genetic structure is shaped by limited dispersal in a naturally fragmented landscape. Ecography 36(6): 731-743.

Bogan, M.T., K.S. Boersma, & D.A. Lytle. Flow intermittency alters longitudinal patterns of invertebrate diversity and assemblage composition in an aridland stream network. Freshwater Biology 58(5): 1016-1028.

Bogan, M.T., OG. Gutierrez, J.A. Alvarado, & D.A. Lytle. Habitat type and permanence determine local aquatic invertebrate community structure in the Madrean Sky Islands. Biodiversity and Management of the Madrean Archipelago III, USDA Forest Service Proceedings RMRS-P-67: 277-282.


McMullen, L.E., & D.A. Lytle. Quantifying invertebrate resistance to floods: a global-scale meta-analysis. Ecological Applications 22(8): 2164-2175.

Daly-Engel, T.S., R.L. Smith, D.S. Finn, M.E. Knoderbane, I.C. Phillipsen, & D.A. Lytle. 17 novel polymorphic microsatellite markers for the giant water bug, Abedus herberti (Belostomatidae). Conservation Genetics Resources: DOI 10.1007/s12686-012-9687-5


Bogan, M.T. & D.A. Lytle. Severe drought and novel community trajectories in desert stream pools. Freshwater Biology 56(10): 270-281.

Konrad, C.P., J.D. Olden, D.A. Lytle, T. Melis, J.C. Schmidt, E. Bray, M. Freeman, K. Gido, N. Hemphill, M. Kennard, L. McMullen, M.C. Mims, M. Pyron, C. Robinson, J.G. Williams. Large scale flow experiments for managing river systems. BioScience 61(12): 948-959.

Larios, N., Lin, J., Zhang, M., Lytle, D., Moldenke, A., Shapiro, L. G., and Dietterich, T. G. Stacked spatial-pyramid kernel: An object-class recognition method to combine scores from random trees. IEEE Workshop on Applications of Computer Vision. DOI: 10.1109/WACV.2011.5711522

Lytle, D.A. Abedus herberti. Pp. 123-126 in Li, J. & M. Barbour (Eds.): Wading for Bugs: Discovering Stream Insects with the Experts. OSU Press.


Lytle, D. A., G. Martínez-Muñoz, W. Zhang, N. Larios, L. Shapiro, R. Paasch, A. Moldenke, E. N. Mortensen, S. Todorovic, T. G. Dietterich. Automated processing and identification of benthic invertebrate samples. J. North American Benthological Association 29(3): 867-874.

Merritt, D. M., Scott, M. L., Poff, N. L., Auble, G. T. & Lytle, D. A. Theory, methods, and tools for determining environmental flows for riparian vegetation: riparian vegetation-flow response guilds. Freshwater Biology 55: 206-225.

Shafroth, P. B., Anderson, D. C., Beauchamp, V. B., Hautzinger, A., Hickey, J. T., Lytle, D. A., McMullen, L. E., Warner, A. & Wilcox, A. C. Models and tools to assist environmental flow assessment and implementation below reservoirs: lessons from an arid-region river. Freshwater Biology 55: 68-85.


Finn, D. S., Bogan, M. T. & Lytle, D. A. Demographic stability metrics for conservation prioritization of isolated populations. Conservation Biology 23(5): 1185-1194.

Martínez-Muñoz, G., W. Zhang, N. Payet, S. Todorovic, N. Larios, A. Yamamuro, D. Lytle, A. Moldenke, E. Mortensen, R. Paasch, L. Shapiro, T. Dietterich. Dictionary-free categorization of very similar objects via stacked evidence trees. Computer Vision and Pattern Recognition 2009: 549-556.

Fox, H., et al. Why fly? Institutional changes to reduce high-flying conservationists' footprints. Frontiers in Ecology & the Environment 7(6): 294-295.


Lytle, D.A. Life-history and behavioural adaptations of aquatic insects in disturbed environments. In J. Lancaster and R. Briers (Eds.) Aquatic Insects: Challenges to Populations. CABI International, London.

Lytle, D.A., J.D. Olden, L.E. McMullen. Drought-escape behaviors of aquatic insects may be adaptations to highly variable flow regimes characteristic of desert rivers. Southwestern Naturalist 53(3): 399-402.

Lytle, D.A., M.T. Bogan, and D.S. Finn. Evolution of aquatic insect behaviors across a gradient of disturbance predictability. Proceedings of the Royal Society - Series B 275: 453-462.

Sarpola, M.J., R.K. Paasch, E.N. Mortensen, T.G. Dietterich, D.A. Lytle, A.R. Moldenke, and L. Shapiro. An aquatic insect imaging system to automate insect classification. Transactions of the American Association of Agricultural and Biological Engineers 51(6): 2217-2225.


Finn, D. S., M. Blouin, D. A. Lytle. Population genetic structure reveals terrestrial affinities for a headwater stream insect. Freshwater Biology 52: 1881-1897.

Bogan, M. T. and D. A. Lytle. Seasonal flow variation allows "time-sharing" by disparate aquatic insect communities in montane desert streams. Freshwater Biology 52: 290-304.

Lytle, D.A., & N. J. White. Rainfall cues and flash-flood escape in desert stream insects. Journal of Insect Behavior 20:413-423.

Hall, S. R., M. A. Leibold, D. A. Lytle, V. Smith. Grazers, producer stoichiometry, and the light:nutrient hypothesis revisited. Ecology 88(5): 1142-1152.

HallEtal2007_Ecol.pdf  (194.76 KB)

Larios, E., H. Deng, W. Zhang, M. Sarpola, J. Yuen, R. Paasch, A. Moldenke, D. A. Lytle, S. R. Correa, E. Mortensen, L. Shapiro, T. Dietterich. Automated insect identification through concatenated histograms of local appearance features: feature vector generation and region detection for deformable objects. Machine Vision and Applications. DOI 10.1007/s00138-007-0086-y.

E. N. Mortensen, E. L. Delgado, H. Deng, D. Lytle, A. Moldenke, R. Paasch, L. Shapiro, P. Wu, W. Zhang, T. G. Dietterich. Pattern Recognition for Ecological Science and Environmental Monitoring: An Initial Report. pp.189-206 in N. MacLeod (Ed.) Automated Taxon Identification in Systematics: Theory, Approaches and Applications. CRC Press, Boca Raton.

Larios, E., H. Deng, W. Zhang, M. Sarpola, J. Yuen, R. Paasch, A. Moldenke, D. A. Lytle, S. R. Correa, E. Mortensen, L. Shapiro, T. Dietterich. Automated insect identification through concatenated histograms of local appearance features. IEEE Workshop on Applications of Computer Vision. DOI 10.1109/WACV.2007.13.


Hall, S. R., M. A. Leibold, D. A. Lytle, V. Smith. Inedible producers in stoichiometrically-explicit food webs: inter-trophic competition and controls on food quality for grazers. American Naturalist 167(5): 626-635.


Hall, S. R., V. Smith, D. A. Lytle, M. A. Leibold. Constraints on primary producer N:P stoichiometry along N:P supply ratio gradients. Ecology 86(7): 1894-1904.


Lytle, D. A. and N. L. Poff. Adaptation to natural flow regimes. Trends in Ecology and Evolution 19(2): 94-100. Assembles known examples of plants and animals that have evolved life history, behavioral, or morphological traits for surviving flood or drought, and discusses what these adaptations mean for stream and river conservation.

Lytle, D. A. and D. M. Merritt. Hydrologic regimes and riparian forests: a structured population model for cottonwood. Ecology 85(9): 2493-2503. Describes a modeling approach that can be used to understand how different cycles of flood and drought affect long-term population dynamics of cottonwood forests.

Lytle, D. A. and R. L. Smith. Exaptation and flash flood escape in the giant water bugs. Journal of Insect Behavior 17(2): 169-178. Describes how flood escape behavior was coopted from an ancient (>150 m.y. old) migratory behavior.

Hall, S. R., V. H. Smith, D. A. Lytle, and M. A. Leibold. Stoichiometry and planktonic grazer composition over gradients of light, nutrients, and predation risk. Ecology 85(8): 2291-2301. Results of a mesocosm experiment that explores species sorting of zooplankton communities across ecological conditions (light, nutrients, predators) observed in natural ponds.


Lytle, D. A. Reconstructing long-term flood regimes with rainfall data: effects of flood timing on caddisfly populations. Southwestern Naturalist 48(1): 36-42. Describes how the statistical properties of monsoon rainstorms can be exploited to estimate flood timing and frequency, and what this means for caddisfly populations.


Lytle, D. A. Flash floods and aquatic insect life-history evolution: evaluation of multiple models. Ecology 83(2): 370-385. A test of theory developed in the 2001 Am. Nat. paper. Caddisfly life histories are temporally synchronized to avoid flash floods.

Lytle2001_Ecology.pdf  (208.86 KB)


Lytle, D. A. Convergent growth regulation in arthropods: biological fact or statistical artifact? Oecologia 128(1): 56-61. Shows how growth increment analysis has undesirable statistical properties - avoid it! Use geometric mean regression instead.

Lytle, D. A. Disturbance regimes and life history evolution. American Naturalist 157(5): 525-536. Describes a model for exploring how disturbance timing, frequency, severity, and predictability affect the evolution of life history strategies. I had aquatic insects in mind, but the theory is general.

Lytle2001_AmNat.pdf  (170.93 KB)

Lytle, D. A. and B. L. Peckarsky. Spatial and temporal impacts of a diesel fuel spill on stream invertebrates. Freshwater Biology 46: 693-704.

Peckarsky, B. L., B. W. Taylor, A. R. McIntosh, M. A. McPeek, & D. A. Lytle. Variation in mayfly size at metamorphosis as a developmental response to risk of predation. Ecology 82(3): 740-757.


Lytle, D. A. Biotic and abiotic effects of flash flooding in a montane desert stream. Archiv für Hydrobiologie 150(1): 85-100.


Lytle, D. A. Use of rainfall cues by Abedus herberti (Hemiptera: Belostomatidae): a mechanism for avoiding flash floods. Journal of Insect Behavior 12(1): 1-12.

Lytle1999_JIB.pdf  (671.22 KB)